Teucrium Cyprium
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''Teucrium'' is a
cosmopolitan Cosmopolitan may refer to: Food and drink * Cosmopolitan (cocktail), also known as a "Cosmo" History * Rootless cosmopolitan, a Soviet derogatory epithet during Joseph Stalin's anti-Semitic campaign of 1949–1953 Hotels and resorts * Cosmopoli ...
genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wid ...
herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in opposite pairs, and flowers arranged in
thyrse A thyrse is a type of inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot o ...
s, the corolla with mostly white to cream-coloured, lobed petals.


Description

Plants in the genus ''Teucrium'' are perennial herbs or shrubs with four-cornered stems, often with simple hairs and
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
glands In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland). Structure De ...
. The leaves are arranged in opposite pairs, simple or with three leaflets sometimes with lobed or serrated edges. The flowers are arranged in a thyrse, sometimes in a cyme in leaf axils. The flowers have five more or less similar
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...
s fused at the base, and the corolla is white or cream-coloured with five lobes forming two lips. The upper lip is usually much reduced in size and the lower lip has three lobes, the central lobe usually larger than the side lobes. There are four stamens attached near the base of the petals and the fruit is a
schizocarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more seeds (the m ...
with four segments.


Taxonomy

The genus ''Teucrium'' was first formally described in 1753 by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...
in '' Species Plantarum''. The name ''Teucrium'' was used by
Pedanius Dioscorides Pedanius Dioscorides ( grc-gre, Πεδάνιος Διοσκουρίδης, ; 40–90 AD), “the father of pharmacognosy”, was a Greek physician, pharmacologist, botanist, and author of '' De materia medica'' (, On Medical Material) —a 5-vo ...
for several species in this genus, and is believed to refer to
King Teucer In Greek mythology, King Teucer (; Ancient Greek: Τεῦκρος ''Teûkros'') was said to have been the son of the river-god Scamander and the nymph Idaea. Mythology Before the arrival of Dardanus, the land that would eventually be called D ...
of
Troy Troy ( el, Τροία and Latin: Troia, Hittite: 𒋫𒊒𒄿𒊭 ''Truwiša'') or Ilion ( el, Ίλιον and Latin: Ilium, Hittite: 𒃾𒇻𒊭 ''Wiluša'') was an ancient city located at Hisarlik in present-day Turkey, south-west of Ç ...
who used the plant in his medicine.


Species

(See also List of ''Teucrium'' species) ''Teucrium'' is a cosmopolitan genus with about 300 species, the distribution centred on the Mediterranean. There are about thirteen species endemic to Australia.


Selected species

* '' Teucrium albicaule'' Toelken – scurfy germander (Australia) * ''
Teucrium arduinoi ''Teucrium'' is a Cosmopolitan distribution, cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial plant, perennial herbaceous plant, herbs or shrubs, with branches that a ...
'' L. * '' Teucrium argutum'' R.Br. – native germander (Qld., N.S.W.) * '' Teucrium aroanium'' Orph. ex Boiss. * '' Teucrium balfourii'' Vierh. * '' Teucrium balthazaris'' Sennen * '' Teucrium betonicum'' L'Hér. * '' Teucrium botrys'' L. – cut-leaved germander * '' Teucrium canadense'' L. – American germander, Canada germander, wood sage, hairy germander * '' Teucrium capitatum'' L. * ''
Teucrium carolipaui ''Teucrium'' is a Cosmopolitan distribution, cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial plant, perennial herbaceous plant, herbs or shrubs, with branches that a ...
'' C. Vicioso ex Pau * ''
Teucrium chamaedrys ''Teucrium chamaedrys'', the wall germander, is a species of plant native to the Mediterranean regions of Europe and North Africa, and the Middle East as far as Iran.Dobignard, A. & Chatelain, C. (2012). Index synonymique de la flore d'Afrique d ...
'' L. – wall germander * '' Teucrium chardonianum'' Maire & Wilczek * '' Teucrium coahuilanum'' B.L.Turner * '' Teucrium corymbosum'' R.Br. – forest germander (Australia, New Guinea) * '' Teucrium cossonii'' D.Wood – fruity teucrium * '' Teucrium creticum'' L. * '' Teucrium cubense'' Jacq. – small coastal germander, dwarf germander * ''
Teucrium eremaeum ''Teucrium eremaeum'' is a species of flowering plant in the family Lamiaceae and is endemic to the south-west of Western Australia. It is a perennial herb or shrub with small, linear to lance-shaped leaves and white or cream-coloured flowers. ...
''
Diels Diels is the last name of several people: * Rudolf Diels (1900–1957), German politician * Otto Diels (1876–1954), German scientist noted for his work on the Diels–Alder reaction * Ludwig Diels (1874–1945), German botanist * Hermann Diels ...
(W.A.) * '' Teucrium divaricatum'' Sieber ex Heldr. * '' Teucrium flavum'' L. * '' Teucrium fruticans'' L. – tree germander, shrubby germander * '' Teucrium glandulosum'' Kellogg – common germander, desert germander, sticky germander * ''
Teucrium gnaphalodes ''Teucrium gnaphalodes'' is a plant species in the genus ''Teucrium''. It is endemic to the Iberian Peninsula and grows at altitudes between 200 and 1500 m. It flowers from March to July. The flavones diosmin, cirsimaritin, salvigenin, cirsili ...
'' L’Hér. * '' Teucrium grandiusculum'' F.Muell. & Tate (W.A., S.A., N.T.) * '' Teucrium heterophyllum'' L’Hér. * '' Teucrium integrifolium'' Benth. – teucry weed (W.A., N.T., Qld.) *'' Teucrium junceum'' * '' Teucrium laciniatum'' Torr. – germander, lacy germander * '' Teucrium lepicephalum'' Pau * '' Teucrium marum'' L. * '' Teucrium massiliense'' L. * ''
Teucrium montanum ''Teucrium montanum'', the mountain germander, is a germander native to southern Europe, from Spain eastward to Turkey. It forms shrubs about a foot high, with small leaves no more than an inch across, and petals blooming yellowish white. On sout ...
'' L. * '' Teucrium oliverianum'' Ging. ex Benth. * '' Teucrium orientale'' L. – oriental germander * '' Teucrium polium'' L. * '' Teucrium pseudochamaepitys'' L. * '' Teucrium puberulum'' * '' Teucrium pyrenaicum'' L. * '' Teucrium racemosum'' – forest germander * ''
Teucrium scordium ''Teucrium scordium'' is a species of flowering plant belonging to the family Lamiaceae. Its native range is Europe to China. References

{{Taxonbar, from=Q158832 Teucrium, scordium Flora of Malta ...
'' L. * ''
Teucrium scorodonia ''Teucrium scorodonia'', common name the woodland germander or wood sage, is a species of flowering plant in the genus ''Teucrium'' of the family Lamiaceae. It is native to Western Europe and Tunisia, but cultivated in many places as an ornament ...
'' L. – woodland germander * '' Teucrium socotranum'' Vierh. * '' Teucrium subspinosum'' Pourr. ex Willd. * '' Teucrium teucriiflorum'' (F.Muell.) Kattari & Salmaki * '' Teucrium townsendii'' ** ''Teucrium townsendii'' ssp. ''affine'' ** ''Teucrium townsendii'' var. ''townsendii'' * '' Teucrium vesicarium'' Mill. * '' Teucrium viscidum'' Blume * '' Teucrium werneri'' Emb.


Fossil record

†''Teucrium tatjanae'' seed
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s are known from the Oligocene,
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
and
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
, Miocene and Pliocene of central and southern
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eig ...
and Miocene of
Lusatia Lusatia (german: Lausitz, pl, Łużyce, hsb, Łužica, dsb, Łužyca, cs, Lužice, la, Lusatia, rarely also referred to as Sorbia) is a historical region in Central Europe, split between Germany and Poland. Lusatia stretches from the Bóbr ...
. The fossil seeds are similar to seeds of the extant ''Teucrium orientale''. †''Teucrium pripiatense'' seed fossils have been described from the Pliocene Borsoni Formation in the Rhön Mountains of central
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
.The floral change in the tertiary of the Rhön mountains (Germany) by Dieter Hans Mai - Acta Paleobotanica 47(1): 135-143, 2007.


References


External links


USDA PLANTS Profile
{{Taxonbar, from=Q157714 * Lamiaceae genera Taxa named by Carl Linnaeus